Coneflowers
Echinacea purpurea Rubinstern - an"improved" purple coneflower seed
strain (also sold as 'Ruby Star,' the English translation
(above) Echinacea pallida (pale coneflower) with
Helianthus slaicifolius (willowleaf sunglower)

(right)
Echinacea tennesseensis (Tennessee coneflower)
with Oputia cactus.
(left) Echinacea paradoxa var. paradoxa (yellow purple coneflower)with Monarda
didyma (bee balm)

(above) Coneflower hybrids and Chicago BOtanic Garden trials.
Echinacea X 'Orange Meadowbright'
Echinacea X 'Sundown'
Echinacea X 'Harvest Moon'
There are very few native plants
with wider appeal than the
coneflower (Echinacea spp). This
uniquely North American genus of
ten species epitomizes the
meadow or prairie wildflower. In
general, coneflowers are easy to
grow in sunny, well-drained soil
and the large daisy-shaped blooms
remain colorful for at least a
month. The plants grow from a
woody crown, sending up first a
tuft of basal leaves followed by
leafy flower stems that rise up in
mid-summer. Each stem is topped
by one large bloom that can be as
much as 4 inches across and once
the first bloom begins to fade
others grow from small side
branches to take its place. Like
other members of the aster family,
Echinacea flowers are technically
inflorescences made up of many
small flowers serving one of two
functions. Each petal or ray is really
a single flower with five petals
fused into one. The ray flowers
form a ring around the central
cone, which is also composed of
many individual flowers that have
no petals. The division of labor has
particle consequences for
pollination and seed set. The rays
are sterile, functioning merely as
advertisement for the less
noticeable fertile flowers that make
up the cone. These later produce
nectar and pollen as well as seeds.
A bee or butterfly cannot help but
notice the ring of large, colorful ray
flowers as it passes, and the cone
provides a perfect lading pad foe the insect as it comes in to investigate. What the bug finds is a host of little, nectar rich blooms
packed in together - a sort of one stop shopping that is very appealing. Rather than wasting energy flitting here and there, the bee or
butterfly can settle in and drink from a bunch of flowers at once. When you plant coneflowers, the butterflies and bees are sure to
follow.
This combination of colorful advertizing
and concentrated, easily accessible flowers
has made the aster family one of the most
successful and diverse in the world.
However, the coneflowers appear to be a
fairly new member of this large family as
most of the species are pretty similar in leaf
and flower shape and color. Coneflowers
likely evolved fairly recently during the
drying of the North American climate
during the Pleistocene that allowed the
development of prairie grassland habitat
from the Southeastern US to the western
mountains. The typical color of the bloom
is light lavender with a deep red or black
cone bristling with yellow or red knobs
(these knobs or bristles likely aid pollination
by providing scaffolding for the pollinator
to crawl on then discourage predation of
seeds by birds).
There are two basic leaf types in the genus - one characterized by a 2-3 inch wide, ovate blade with a rounded or heart-shaped base
and the other by a narrow (<1 inch) wide) linear leaf. The narrow-leaved species tend to grow in drier habitats than the two
wide-leaved ones (E. purpurea - purple coneflower, which is found in the prairies west of the Appalachians and E. laevigata - smooth
purple coneflower, which is a rare species from the eastern side of those same mountains). The most widespread of the narrow-leaved
species is E. pallida (pale coneflower from the prairies east of the Rocky Mountains. It is closely related to the smaller E. angustifolia
(narrow-leaved purple coneflower) which grows on both sides of the Rockies. There are several other related species, notable E.
tennesseensis (Tennessee coneflower), which is very similar to E. angustifolia but restricted to dry, rocky prairie remnants on the
Cumberland Plateau of Tennessee. Like most coneflowers, it is adapted to rather parched, fire-prone grasslands. From a horticultural
point of view this is important as most of the species resent wet soils and poor air circulation around the leaves and crown of the
plant. They have large, fleshy taproots that burrow deep in search of moisture and act as a food pantry to help the plant recover after
drought or the frequent fires that sweep through their habitat. Taproots resent disturbance, rot easily if drainage is poor, and generally
make plants more difficult to transplant and cultivate. The exception in this genus is E. purpurea, which prefers damp or even wet
prairies and as a consequence has evolved a more forgiving fibrous root system that functions much better in this type of soil. From a
functional point of view, purple coneflower is a much easier and forgiving plant to cultivate and it is no wonder that it has become the
unquestionable favorite among gardeners. It has a further advantage in that its ray petals are wide and flat unlike the narrow or curled
rays of the other species. The rays of wild plants point downward much like the feathers of a badminton shuttlecock, though breeders
have selected forms which hold their rays parallel to the ground for flatter appearance. Tennessee coneflower is the only species with
rays that naturally orient lightly upwards, and breeders have crossed this one with purple coneflower to yield hybrids with even flatter
blooms.